一、发表学术论文

[1] Liu L^#, Cao X^#, Zhai Z, Ma S, Tian Y, Cheng JL* (2022) Direct evidence of drought stress memory in mulberry from a physiological perspective: Antioxidative, osmotic and phytohormonal regulations, Plant Physiology and Biochemistry, 186, 76-87.

[2] Cao X, Shen Q, Ma S, Liu L, Cheng JL^*(2020) Physiological and PIPtranscriptional responses to progressive soil water deficit in three mulberry cultivars. Frontiers in Plant Science, 11, 1310.

[3] Cao X, Shen Q, Liu L, Cheng JL^* (2020) Relationships of growth, stable carbon isotope composition and anatomical properties of leaf and xylem in seven mulberry cultivars: a hint towards drought tolerance. Plant Biology, 22,287-297.

[4] Cao X, Yang H, Shang C, Ma S, Liu L, Cheng JL^* (2019) The roles of auxin biosynthesis YUCCA gene family in plants. International Journal of Molecular Sciences, 20, 6343.

[5] Shang C, Yang H, Ma S, Shen Q, Liu L, Hou C, Cao X^*, Cheng JL^* (2019) Physiological and transcriptomic changes during the early phases of adventitious root formation in mulberry stem hardwood cuttings. International Journal of Molecular Sciences, 20, 3707.

[6] Cao X, Shen Q, Shang C, Yang H, Liu L, Cheng JL^* (2019) Determinants of shoot biomass production in mulberry: combined selection with leaf morphological and physiological traits. Plants, 8, 118.

[7] Cao X^#, Du W^#, Shang CQ, Shen QD, Liu L, Cheng JL^* (2018) Comparative transcriptome reveals circadian and hormonal control of adventitious rooting in mulberry. Acta Physiologiae Plantarum, 40(11):197.

[8] Du XL^#, Cao X^#, Yin CR, Tang Z, Du W, Ban YY, Cheng JL^* (2017) Comprehensive analysis of R2R3-MYB genes during adventitious root formation in cuttings of Morus alba. Journal of Plant Growth Regulation 36: 290-299.

[9] Cao X, Jia J, Zhang C, Li H, Liu TX, Jiang X, Polle A, Peng CH, Luo ZB^* (2014) Anatomical, physiological and transcriptional responses of two contrasting poplar genotypes to drought and re-watering. Physiologia Plantarum 151: 480-494.

[10] Cao X, Jia J, Li H, Li MC, Luo J, Liang ZS, Liu TX, Liu WG, Peng CH, Luo ZB^* (2012) Photosynthesis, water use efficiency and stable carbon isotope composition are associated with anatomical properties of leaf and xylem in six poplar species. Plant Biology 14: 612-620.

[11] Chao N, Chen WQ, Cao X, JiangXN, GaiY, Liu L^* (2022)  Plant hormones coordinate monolignol biosynthesis with seasonal changes in Populus tomentosa, Environmental and Experimental Botany, 195, 104784.

[12] Liu L, Chao N, Yidilisi K, Kang X, Cao X (2022) Comprehensive analysis of the MYB transcription factor gene family in Morus alba, BMC Plant Biology, 22, 281.

[13] Jia J, Zhou J, Shi W, Cao X, Luo J, Polle A, Luo ZB^* (2017) Comparative transcriptomic analysis reveals the roles of overlapping heat-/drought-responsive genes in poplars exposed to high temperature and drought. Scientific Reports 7: 43215.

[14] Jia J, Li SJ, Cao X, Li H, Shi WG, Polle A, Liu TX, Peng CH, Luo ZB^* (2016) Physiological and transcriptional regulation in poplar roots and leaves during acclimation to high temperature and drought. Physiologia Plantarum 157: 38-53.

[15] Li H, Li MC, Luo J, Cao X, Qu L, Gai Y, Jiang X, Liu TX, Bai H, Janz D, Polle A, Peng CH, Luo ZB^* (2012) N-fertilization has different effects on the growth, carbon and nitrogen physiology, and wood properties of slow- and fast-growing Populus species. Journal of Experimental Botany 63: 6173-6185.

[16] 陶然，曹旭*，吴华顺，杨洪雷，马桑，刘利，程嘉翎*.干旱胁迫后叶面追施钼盐对桑树生长和部分生理指标的影响[J].蚕业科学,2020,46(05):529-535.

[17]杨洪雷，吴华顺，马桑，尚春琼，刘利，曹旭*，程嘉翎*.叶面喷施锌和钼盐对干旱胁迫下桑树生长和部分生理指标的影响[J].蚕业科学，2020，46(04)：418-425.

[18] 杜伟，曹旭，黄平，程嘉翎*. 9个基因在桑树硬枝扦插生根过程中的表达特性分析[J].蚕业科学，2019，45(02)：157-164.

[19] 沈萩荻，吴艳，刘利，曹旭*，程嘉翎*. 六个不同品种桑树叶柄解剖结构与水分运输能力初探[J]. 蚕业科学，2019，45（1）：0001-0008.

[20] 殷朝瑞，方荣俊，尚春琼，沈萩荻，曹旭，程嘉翎. 3个实用桑树品种的耐盐性生理生化特征及耐盐害的能力评价[J]. 蚕业科学，2018，44（3）: 0359-0366.  

[21] 杜小龙，曹旭，刘利，殷朝瑞，班月圆，李建龙，程嘉翎. 转录组与蛋白质组关联分析鉴定桑树硬枝扦插皮部生根相关功能基因[J]. 蚕业科学，2017，43（03）：374~381.

[22] 杜伟，曹旭，殷朝瑞，方荣俊，郭锡杰，刘利，程嘉翎. 桑树硬枝扦插不同生根类型插穗的生理特性研究[J]. 蚕业科学，2017，43（01）：18~24.

[23] 班月圆，方荣俊，杜伟，殷朝瑞，曹旭，刘利，程嘉翎，盐胁迫下3个地域来源桑种子的萌发率及部分生化性状差异[J]. 蚕业科学，2016，42（6）：960~967

二、教研论文

曹旭.《植物生理学实验》课程教学改革的实践与思考 [J].教育现代化，2019，6(66)：113-114+130.

三、参编论著

中国桑树栽培学.中国农业科学院蚕业研究所 江苏科技大学 主编.上海科学技术出版社.2020